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Movements and Habitat Selection by Raccoons in a Fragmented Agricultural
Landscape in Illinois
Erin Barding and Thomas Nelson
Department of Biological Sciences, Eastern Illinois
University, Charleston, IL
Introduction
Raccoons are found throughout most of North America, occupying a
wide variety of habitats from swamps to prairies, farms and cities. During the
past twenty years, populations have increased dramatically in Illinois due in
part to low pelt prices and reduced hunting pressure (Gehrt et al. 2002).
Illinois’ rural landscape also supports high raccoon densities because corn
fields provide important supplemental food during winter. On a gradient from
forested to cultivated, raccoon densities tend to be highest in mid- to highly
cultivated areas (Pedlar et al. 1997).
At these densities, raccoons may exert levels of predation rarely seen in
natural settings. For example, extremely high rates of nest predation have been
reported near forest edges in Illinois (Robinson 1992). A recent study reported
that “raccoon-vulnerable” birds (those nesting below eight feet) have declined
during the past twenty years, whereas “raccoon-invulnerable” birds increased
(Schmidt 2003). Collectively, these studies suggest that high raccoon densities
may be an important factor influencing the abundance and diversity of songbirds
in Illinois.
Numerous studies have reported generalized habitat use by raccoons, but few have
been conducted in highly fragmented, agricultural landscapes or focused on
habitat selection by raccoons during the avian nesting season. We investigated
summer movements and habitat selection by raccoons occupying a rural Illinois
landscape at two spatial scales, the home range and habitat types within the
home range. This information may help us identify those habitat complexes most
vulnerable to raccoon predation.
Objectives
1. Evaluate the movements and home ranges of raccoons during the
avian nesting and brood rearing season.
2. Describe patterns of habitat selection at two spatial scales during the
summer.
Methods
Research was conducted on the Richardson Wildlife Foundation (RWF)
property in northern Illinois. This 1,000 ha site is composed of wetlands (10%),
restored prairies (28%), forests (37%) and crops (22%). Raccoons were
live-trapped, sexed, aged, and radio-collared. Collared raccoons were located
three to four time each week from May to September. Individual animals were
tracked during continuous four-hour intervals at night to investigate movements
and habitat use while foraging. Locations during these periods was recorded
every fifteen minutes and recorded in
Universal Transverse Mercator (UTM) coordinates. We
used the Animal Movement extension and ArcView software to estimate
minimum convex polygon (MCP) and 95% kernel home
ranges. Home ranges were intersected with a GIS habitat map of the study area to
estimate the habitat composition of each range. Habitats were classified as
forest, forest edge, wetland, grassland, crops or residential
(Figure 1). We assessed habitat use at two scales using
compositional analysis (Aebischer et al. 1993). We first compared the
habitat composition of home ranges to the composition of the study area. We then
compared habitat associated with raccoon locations to habitat composition of the
home range. When overall use of habitat was non-random, we tested all possible
habitat pairs and ranked them according to relative use.
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Figure 1. Example of MCP,
95% kernel and 50% kernel for a raccoon radio-tracked on RWF during
summer 2002. |
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Figure 2. Mean habitat availability
(study area) and use (within study areas and radio locations) by
raccoons during summer. |
Results
The median MCP summer range was xx.x ha (range: ) and did
not differ between males and females (U = 26.0; P = 0.09).
Median core area (50% kernel) was xx.x ha (range: ) and did not differ between
the sexes
(U = 37.0; P = 0.37).
During foraging bouts males averaged xx.x m/hr; whereas females moved less,
averaging xx.x m/hr
(U = 4.0; P = 0.04).
Proportions of habitats within home ranges were different than proportions
available in the study area (Λ = 0.26, P < 0.01). Home ranges contained more
forest edge and less cropland than study areas (Figure 2).
Proportions of raccoon locations within habitats differed from proportions
within home ranges
(Λ = 0.04; P < 0.01). Within their home ranges, raccoons
selected forest edges and wetlands for foraging and avoided grasslands and
cropland (Figure 2).
Based on relative use of habitats within home ranges, foraging
habitats rank in order of preference as forest edge, wetlands, residential,
forest interiors, grasslands and cropland.
Conclusions
Raccoons foraged predominantly in forest edges and wetlands
during the summer, but grasslands were used infrequently. Cornfields were
avoided early in the summer when they provided little cover, but use increased
as these fields matured in late summer. Forest/cropfield and forest/grassland
edges were used extensively as travel lanes. If raccoons predate nests
incidentally while foraging on other foods, nest predation on grassland birds
may be reduced by maintaining or restoring large grassland tracts and separating
nesting habitat from raccoon travel lanes and foraging areas.
Literature Cited
Aebisher, N., P. Robertson, and R. Kenward. 1993. Compositional
analysis of habitat use from animal radio-tracking data. Ecology 74:1313-1325.
Gehrt, S., G. Hubert, Jr., and J. Ellis. 2002. Long-term population trends of
raccoons in Illinois. Wildlife Society Bulletin 30:457-463.
Pedlar, J., L. Fahrig, and H. Merriam. 1997. Raccoon habitat use at 2 spatial
scales. J. Wildlife. Management. 61:102-112.
Robinson, S., F. Thompson III., T. Donovan, D. Whitehead, and J. Faaborg. 1995.
Regional forest fragmentation and the nesting success of migratory birds.
Science 267:1987-1990.
Schmidt, K. 2003. Nest predation and population declines in Illinois songbirds:
a case for mesopredator effects. Conservation Biology 17:1141
Acknowledgements
This research is funded by the Illinois Natural History Survey (INHS) and
Eastern Illinois University.
Thanks to J. Crawford, Dr. P. Brown (INHS) and T. Moyer (RWF) for assistance and
Richardson’s Wildlife Foundation for providing housing and access to the study
area.
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